Course Summary Course Objectives Learning Outcomes Course Materials Teaching Methods Weekly Topics Course Schedule Office Hours Assestment ECTS Calculation Course Policies Learning Tips Print Syllabi Download as PNG

BIO416 Population Genetics

Syllabus   |  International University of Sarajevo  -  Last Update on Mar 03, 2026

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Genetics and Bioengineering

Spring 2023 - 2024 | 6 ECTS Credits | International University of Sarajevo

Academic Year
2023 - 2024
Semester
Spring
Course Code
BIO416
Weekly Hours
3 Teaching + 0 Practice
ECTS
6
Prerequisites
Junior Standing
Teaching Mode Delivery
Face-to-face
Prerequisite For
-
Teaching Mode Delivery Notes
-
Cycle
I Cycle
Prof. Jane Doe

Muhamed Adilović

Course Lecturer

Position
Assistant Professor Dr.
Email
madilovic@ius.edu.ba
Phone
033 957 219
Assistant(s)
-
Assistant E-mail
-

Course Objectives

Introduce students to Population Genetics, focusing on main concepts including Hardy-Weinberg equilibrium, inbreeding, mutations, drift, selection, gene flow, and human population structure.

Learning Outcomes

After successful completion of the course, the student will be able to:

1
Apply the principles of Hardy-Weinberg Equilibrium and model allele, genotype, and phenotype frequencies.
2
Model how the major evolutionary forces—mutation, genetic drift, natural selection, and gene flow—act individually and in concert to alter allele frequencies in populations over time.
3
Analyze the effects of non-random mating, specifically inbreeding, on population structure and genetic diversity.
4
Integrate concepts of gene flow and genetic drift to explain the development of human population structure and history.
5
Develop and test hypotheses about the action of natural selection in shaping human genetic variation and adaptation.

Course Materials

Required Textbook

Human Population Genetics by John H. Relethford

Additional Literature
A Primer of Population Genetics and Genomics by Daniel L. Hartl Molecular population genetics by Hahn, Matthew William

Teaching Methods

Lectures
Presentations
Class Discussions
Problem-solving
Project.

Weekly Topics

This weekly planning is subject to change with advance notice.
Week Topic Readings / References
1 Introduction to the Course
2 Genetic, Mathematical, and Anthropological Background Chapter 1
3 Hardy–Weinberg Equilibrium Chapter 2
4 Inbreeding Chapter 3
5 Mutation; Quiz Chapter 4
6 Problem solving
7 Genetic Drift Chapter 5
8 Models of Natural Selection Chapter 6
9 Midterm
10 Natural Selection in Human Populations Chapter 7
11 Gene Flow Chapter 8
12 Project Preparation and Discussion
13 Human Population Structure and History; Quiz Chapter 9
14 Study of Quantitative Traits Hartl, Chapter 8-9
15 Review

Course Schedule (All Sections)

SectionTypeDay 1Venue 1Day 2Venue 2
BIO416.1 Course - - - -

Office Hours & Room

DayTimeOfficeNotes
Tuesday 08:00 - 11:00 A F1.33
Wednesday 08:00 - 11:00 A F1.33

Assessment Methods and Criteria

Assessment Components

40%x1
Final Exam
AI: Not Allowed

Alignment with Learning Outcomes :  1  2  3  4  5

21%x1
Midterm
AI: Not Allowed

Alignment with Learning Outcomes :  1  2  3  4  5

18%x2
Quizes
AI: Not Allowed

Alignment with Learning Outcomes :  1  2  3  4  5

21%x1
Project
AI: Not Allowed

Alignment with Learning Outcomes :  1  2  3  4  5

IUS Grading System

Grading Scale IUS Grading System IUS Coeff. Letter (B&H) Numerical (B&H)
0 - 44 F 0 F 5
45 - 54 E 1
55 - 64 C 2 E 6
65 - 69 C+ 2.3 D 7
70 -74 B- 2.7
75 - 79 B 3 C 8
80 - 84 B+ 3.3
85 - 94 A- 3.7 B 9
95 - 100 A 4 A 10

IUS Grading System

Letter marks that do not affect student's CGPA:
  • "IP" – In progress is assigned for recording unfulfilled student obligations related to graduation project/thesis/dissertation and internship.
  • "S" – Satisfactory is assigned to a student who passed the examinations that are not numerically graded or whose written assignment has been accepted.
  • "U" – Unsatisfactory is assigned to a student who failed to pass the examinations that are not numerically graded.
  • "W" – Withdrawal signifies that student has withdrawn from the relevant course.
Additional letter mark that affects student's CGPA:

"N/A" – Not attending, and it is assigned to a student who is suspended from the course or who does not meet the minimal requirement for attendance on lectures or tutorials. The course lecturer must follow the attendance policy and assign "N/A" in each case of a student failing attendance.

Late Work Policy

Information about late submission policies will be shared during class and posted in this section. Please check back for official guidelines.

ECTS Credit Calculation

📚 Student Workload

This 6 ECTS credit course corresponds to 150 hours of total student workload, distributed as follows:

Lectures

45 hours ⏳ (15 week × 3 h)

Quizes

10 hours ⏳ (2 week × 5 h)

Midterm

10 hours ⏳ (1 week × 10 h)

Project

20 hours ⏳ (2 week × 10 h)

Final

20 hours ⏳ (2 week × 10 h)

Home study

45 hours ⏳ (15 week × 3 h)

150 Total Workload Hours

6 ECTS Credits


Course Policies

Academic Integrity

All work submitted must be your own. Plagiarism, cheating, or any form of academic dishonesty will result in disciplinary action according to university policies. When in doubt about citation practices, consult the instructor.

Attendance Policy

Students are expected to adhere to the attendance requirements as outlined in the International University of Sarajevo Study Rules and Regulations. Excessive absences, whether excused or unexcused, may impact academic performance and eligibility for assessment. Mandatory sessions (e.g., labs, workshops) require attendance unless formally exempted. For detailed policies on absences, documentation, and penalties, please refer to the official university regulations.

Technology & AI Policy

Laptops/tablets may be used for note-taking only during lectures. Phones should be silenced and put away during all class sessions. Audio/video recording requires prior permission from the instructor.

Artificial Intelligence (AI) Usage: The use of AI tools (e.g., ChatGPT, Copilot, Gemini) varies by assessment component. Please refer to the AI usage indicator next to each assessment item in the Assessment Methods and Criteria section above. Submitting AI-generated content as your own work, where AI is not explicitly allowed, constitutes an academic integrity violation.

Communication Policy

All course-related communication should occur through official university channels (institutional email or SIS). Emails should include [BIO416] in the subject line.

Academic Quality Assurance Policy

Course Academic Quality Assurance is achieved through Semester Student Survey. At the end of each academic year, the institution of higher education is obliged to evaluate work of the academic staff, or the success of realization of the curricula.

More info

Article 112: Evaluation of Work of the Academic Staff

  1. At the end of each academic year, the institution of higher education is obliged to evaluate work of the academic staff, or the success of realization of the curricula.
  2. Evaluation of work of each academic staff member is to be carried out in accordance with the Statute of the institution of higher education by the institution as well as by students.
  3. The institutions of higher education are obliged to carry out a students’ evaluation survey on the academic staff performance after the end of each semester, or after the completed teaching cycle for the subject taught.
  4. Evaluation must evaluate: lecture quality, student-academic staff interaction, correctness of communication, teacher’s attitudes towards students attending the teaching activities and at assessments, availability of suggested reading material, attendance and punctuality of the teacher, along with other criteria which are defined in the Statute.
  5. The institution of higher education by a specific act determines the procedure for evaluation of the academic staff performance, the content of survey forms, the manner of conducting the evaluation, grading criteria for the evaluation, as well as adequate measures for the academic staff who received negative evaluation for two consecutive years.
  6. The evaluation of the academic staff performance is an integral process of establishment the quality assurance system, or self-control and internal quality assurance.
  7. Results of the evaluation of the academic staff performance are to be adequately analyzed by the institution of higher education, and the decision of the head of the organizational unit about the employee’s work performance is an integral part of the personal file of each member of academic staff.

Learning Tips

Engage Actively

Be prepared to contribute thoughtfully during class discussions, labs, or collaborative work. Active participation deepens understanding and encourages critical thinking.

Read and Review Purposefully

Complete assigned readings or prep materials before class. Take notes, highlight key ideas, and jot down questions. Aim to grasp core concepts and their applications—not just facts.

Think Critically in Assignments

Use course frameworks or methodologies to analyze problems, case studies, or projects. Begin early to allow time for reflection and refinement. Seek feedback to improve your work.

Ask Questions Early

Don’t hesitate to reach out when something is unclear. Use office hours, discussion boards, or peer networks to clarify concepts and stay on track.

Course Academic Quality Assurance: Semester Student Survey

Syllabus Last Updated on Mar 03, 2026 | International University of Sarajevo

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Referencing Curricula Print this page

Course Code Course Title Weekly Hours* ECTS Weekly Class Schedule
T P
BIO416 Population Genetics 3 0 6 Tuesday 09:00-11:50
Prerequisite Junior Standing It is a prerequisite to -
Lecturer Muhamed Adilović Office Hours / Room / Phone
Tuesday:
8:00-11:00
Wednesday:
8:00-11:00
A F1.33 - 033 957 219
E-mail madilovic@ius.edu.ba
Assistant Assistant E-mail
Course Objectives Introduce students to Population Genetics, focusing on main concepts including Hardy-Weinberg equilibrium, inbreeding, mutations, drift, selection, gene flow, and human population structure.
Textbook Human Population Genetics by John H. Relethford
Additional Literature
  • A Primer of Population Genetics and Genomics by Daniel L. Hartl
  • Molecular population genetics by Hahn, Matthew William
Learning Outcomes After successful  completion of the course, the student will be able to:
  1. Apply the principles of Hardy-Weinberg Equilibrium and model allele, genotype, and phenotype frequencies.
  2. Model how the major evolutionary forces—mutation, genetic drift, natural selection, and gene flow—act individually and in concert to alter allele frequencies in populations over time.
  3. Analyze the effects of non-random mating, specifically inbreeding, on population structure and genetic diversity.
  4. Integrate concepts of gene flow and genetic drift to explain the development of human population structure and history.
  5. Develop and test hypotheses about the action of natural selection in shaping human genetic variation and adaptation.
Teaching Methods Lectures, Presentations, Class Discussions, Problem-solving, Project.
Teaching Method Delivery Face-to-face Teaching Method Delivery Notes
WEEK TOPIC REFERENCE
Week 1 Introduction to the Course
Week 2 Genetic, Mathematical, and Anthropological Background Chapter 1
Week 3 Hardy–Weinberg Equilibrium Chapter 2
Week 4 Inbreeding Chapter 3
Week 5 Mutation; Quiz Chapter 4
Week 6 Problem solving
Week 7 Genetic Drift Chapter 5
Week 8 Models of Natural Selection Chapter 6
Week 9 Midterm
Week 10 Natural Selection in Human Populations Chapter 7
Week 11 Gene Flow Chapter 8
Week 12 Project Preparation and Discussion
Week 13 Human Population Structure and History; Quiz Chapter 9
Week 14 Study of Quantitative Traits Hartl, Chapter 8-9
Week 15 Review
Assessment Methods and Criteria Evaluation Tool Quantity Weight Alignment with LOs AI Usage
Final Exam 1 40 1,2,3,4,5 Not Allowed
Semester Evaluation Components
Midterm 1 21 1,2,3,4,5 Not Allowed
Quizes 2 18 1,2,3,4,5 Not Allowed
Project 1 21 1,2,3,4,5 Not Allowed
***     ECTS Credit Calculation     ***
 Activity Hours Weeks Student Workload Hours Activity Hours Weeks Student Workload Hours
Lectures 3 15 45 Quizes 5 2 10
Midterm 10 1 10 Project 10 2 20
Final 10 2 20 Home study 3 15 45
        Total Workload Hours = 150
*T= Teaching, P= Practice ECTS Credit = 6
Course Academic Quality Assurance: Semester Student Survey Last Update Date: 27/03/2026

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